ARCHIVES WPL CL Papilionoidea

ARCHIVES WPL CL Papilionoidea
CHECKLIST PAPILIONOIDEA W-PAL

A dynamic checklist of the Western Palearctic butterflies hyperlinked to the original descriptions
at species, genus and family level
(Lepidoptera, Papilionoidea)

Michel Taymans

Sylvain Cuvelier

27.v.2024


Introduction	Papilionoidea	Papilionidae	Hesperiidae	Pieridae	Riodinidae	Lycaenidae	Nymphalidae	Sources	Thx	Updates

Status

Genitalia

BOLD

Range

Synonyms

G♂ G♀

Order Lepidoptera Linnaeus, 1758

ORIG: Lepidoptera Linnaeus, 1758. Syst. Nat. (ed. 10)1: 341, 343, 458.

Suborder Glossata Fabricius, 1775

ORIG: Glossata Fabricius, 1775. Syst. entomol.: 442.

Superfamily Papilionoidea Latreille, [1802]

ORIG: Papilionides Latreille, [1802]. Hist. Nat. Crust. Ins. 3: 387.
TG: Papilio Linnaeus, 1758.

oo

Family Hesperiidae Latreille, 1809

ORIG: Hesperides Latreille, 1809. Gen. Crust. Ins. 4: 187, 207.
TG: Hesperia Fabricius, 1793.

Subfamily Heteropterinae Aurivillius, 1925

ORIG: Heteropterinae Aurivillius, 1925. In: Seitz, Gross-Schmet. Erde 13: 506, 546.
TG: Heteropterus Duméril, 1806.

Tribe Heteropterini Aurivillius, 1925

ORIG: Heteropterinae Aurivillius, 1925. In: Seitz, Gross-Schmet. Erde 13: 506, 546.
TG: Heteropterus Duméril, 1806.

Genus Heteropterus Duméril, 1806

ORIG: Heteropterus Duméril, 1806. Zool. analyt.: 271.
TS: Papilio aracinthus Fabricius, 1777 (= Papilio morpheus Pallas, 1771)

Heteropterus morpheus (Pallas, 1771)

ORIG: Papilio morpheus Pallas, 1771. Reise verschied. Prov. russ. Reichs 1: 471.
TL: Samara region (Russia Federation).

Genus Carterocephalus Lederer, 1853

ORIG: Carterocephalus Lederer, 1853. Verh. zool.-bot. Ver. Wien 2 (Abhandl.): 26.
TS: Papilio paniscus Fabricius, 1775 (= Papilio palaemon Pallas, 1771).

Carterocephalus silvicola (Meigen, [1829])

ORIG: Hesperia silvicola Meigen, [1829]. Syst. Beschr. eur. Schmett. 2: 65-66, pl. 55, fig. 7a-d.
TL: Braunschweig (Lower Saxony, Germany).

Carterocephalus palaemon (Pallas, 1771)

ORIG: Papilio palaemon Pallas, 1771. Reise verschied. Prov. russ. Reichs 1: 471.
TL: Volga region (Russian Federation).

Subfamily Hesperiinae Latreille, 1809

ORIG: Hesperides Latreille, 1809.Gen. Crust. Ins.4: 187, 207.
TG: Hesperia Fabricius, 1793.

Tribe Baorini Doherty, 1886

ORIG: Baorinae Doherty, 1886. J. asiat. Soc. Bengal. 55(2): 111.
TG: Baoris Moore, 1881 [Indomalayan fauna].

Genus Pelopidas Walker, 1870

ORIG: Pelopidas Walker, 1870. Entomologist 5(76): 56.
TS: Pelopidas midea Walker, 1870 (= Gegenes thrax Hübner, [1821]).

Pelopidas thrax (Hübner, [1821])

ORIG: Gegenes thrax Hübner, [1821]. Samml. exot. Schmett. 2: pl. [150], fig. 1-4.
TL: Cairo (Egypt).

Genus Borbo Evans, 1949

ORIG: Borbo Evans, 1949. Catal. Hesp. Eur. Asia Austr. Brit. Mus.: 44, 436.
TS: Hesperia borbonica Boisduval, 1833.

Borbo borbonica (Boisduval, 1833)

ORIG: Hesperia borbonica Boisduval, 1833. Nouv. Ann. Mus. Hist. nat. 2(2): 213-214.
TL: Réunion (Overseas department, France).

Genus Gegenes Hübner, [1819]

ORIG: Gegenes Hübner, [1819]. Verz. bekannt. Schmett.: 107.
TS: Papilio pumilio Hoffman[n]segg, 1804.

Gegenes pumilio (Hoffman[n]segg, 1804)

ORIG: Papilio pumilio Hoffman[n]segg, 1804. Mag. Insektenk. 3: 202.
TL: Amalfi (Salerno prov., Italy).

Gegenes nostrodamus (Fabricius, 1793)

ORIG: Hesperia nostrodamus Fabricius, 1793. Entomol. Syst. 3(1): 328-329.
TL: NW Africa (Algeria).

Tribe Hesperiini Latreille, 1809

ORIG: Hesperides Latreille, 1809. Gen. Crust. Ins. 4: 187, 207.
TG: Hesperia Fabricius, 1793.

Genus Ochlodes Scudder, 1872

ORIG: Ochlodes Scudder, 1872. Report Peabody Acad. Sci. 1872: 57.
TS: Hesperia nemorum Boisduval, 1852 (= Hesperia agricola Boisduval, 1852) [Nearctic species].

Ochlodes sylvanus (Esper, 1777)

ORIG: Papilio sylvanus Esper, 1777. Schmett. Abb. Nat. 1(1): pl. 36 (suppl. 12), fig. 1 ; [1779]: 343.
TL: Franconia (N Bavaria, Germany).

Genus Hesperia Fabricius, 1793

ORIG: Hesperia Fabricius, 1793. Entomol. Syst. 3(1): 258.
TS: Papilio comma Linnaeus, 1758.

Hesperia comma (Linnaeus, 1758)

ORIG: Papilio comma Linnaeus, 1758. Syst. Nat. (ed. 10)1: 484.
TL: Sweden.

Tribe Thymelicini Tutt, 1906

ORIG: Thymelicinae Tutt, 1906. Nat. Hist. Brit. Lepid. 8: 90.
TG: Thymelicus Hübner, [1819].

Genus Thymelicus Hübner, [1819]

ORIG: Thymelicus Hübner, [1819]. Verz. bekannt. Schmett.: 113.
TS: Papilio acteon Rottemburg, 1775.

Thymelicus christi Rebel, 1894

ORIG: Thymelicus christi Rebel, 1894. Ann. Naturhist. Hofmus. 9(1): 41-42, pl. 1, fig. 2.
TL: Tenerife (Canary Islands, Spain).

Thymelicus acteon (Rottemburg, 1775)

ORIG: Papilio acteon Rottemburg, 1775. Naturforscher 6: 30-31.
TL: Landsberg an der Warthe, current city of Gorzów Wielkopolski (Lubuskie prov., Poland).

Thymelicus hamza (Oberthür, 1876)

ORIG: Hesperia hamza Oberthür, 1876. Étud. Entomol. 1: 28-29, pl. 3, fig. 2a-c.
TL: Oran (Algeria).

Thymelicus hyrax (Lederer, 1861)

ORIG: Hesperia hyrax Lederer, 1861. Wiener entomol. Monatschr. 5(5): 149-150, pl. 1, fig. 6.
TL: Antioch, current city of Antakya (Hatay prov., Turkey).

Thymelicus sylvestris (Poda, 1761)

ORIG: Papilio sylvestris Poda, 1761. Ins. Mus. Graec.: 79.
TL: Graz (Austria).

Thymelicus lineola (Ochsenheimer, 1808)

ORIG: Papilio lineola Ochsenheimer, 1808. Schmett. Eur. 1(2): 230-231.
TL: Germany.

Subfamily Pyrginae Burmeister, 1878

ORIG: Pyrgidae Burmeister, 1878. Descr. phys. Rép. Argentine 5: 245-246.
TG: Pyrgus Hübner, [1819].

Tribe Carcharodini Verity, 1940

ORIG: Carcharodidi Verity, 1940. Farfalle diurne Italia 1: 10.
TG: Carcharodus Hübner, [1819].

Genus Spialia Swinhoe, [1912]

ORIG: Spialia Swinhoe, [1912]. Lepid. Indica 10: 99.
TS: Hesperia galba Fabricius, 1793 [S Asia fauna].

Subgenus Spialia Swinhoe, [1912]

ORIG: Spialia Swinhoe, [1912]. Lepid. Indica 10: 99.
TS: Hesperia galba Fabricius, 1793 [S Asia fauna].

Spialia ali (Oberthür, 1881)

ORIG: Syrichthus ali Oberthür, 1881. Étud. Entomol. 6: 61, pl. 2, fig. 3.
TL: Oran (Algeria).

Spialia therapne (Rambur, 1832)

ORIG: Hesperia therapne Rambur, 1832. Ann. Soc. entomol. Fr. 1: 265, pl. 7, fig. 4.
TL: Corsica (S France).

Spialia sertorius (Hoffman[n]segg, 1804)

ORIG: Papilio sertorius Hoffman[n]segg, 1804. Mag. Insektenk. 3: 203.
TL: Germany.

Spialia rosae Hernández-Roldán, Dapporto, Dincă, Vicente & Vila, 2016

ORIG: Spialia rosae Hernández-Roldán, Dapporto, Dincă, Vicente & Vila, 2016. Molecular Ecology 25(17): 4281 (p. 15).
TL: Puerto de la Ragua (Sierra Nevada, Granada prov., Spain).

Spialia orbifer (Hübner, [1823])

ORIG: Papilio orbifer Hübner, [1823]. Samml. eur. Schmett. 1: pl. 161, fig. 803-806.
TL: Hungary.

Subgenus Platygnathia Picard, [1948]

ORIG: Platygnathia Picard, [1948]. Bull. Soc. entomol. Fr. 52(8): 132.
TS: Hesperia phlomidis Herrich-Schäffer, 1845.

Spialia phlomidis (Herrich-Schäffer, [1845])

ORIG: Hesperia phlomidis Herrich-Schäffer, [1845]. Syst. Bearb. Schmett. Eur. 1: 153, 164, pl. (Hesperides) 2, fig. 8-9.
TL: Sea of Marmara (NW Turkey).

Spialia doris (Walker, 1870)

ORIG: Nisoniades doris Walker, 1870. Entomologist 5(76): 56-57.
TL: Tajora (Bab-el-Mandeb Strait, Djibouti).

Genus Favria Tutt, 1906

ORIG: Favria Tutt, 1906. Nat. Hist. Brit. Lepid. 8: 218.
TS: Hesperia cribrellum Eversmann, 1841.

Favria cribrellum (Eversmann, 1841)

ORIG: Hesperia cribrellum Eversmann, 1841. Bull. Soc. imp. nat. Mosc. 1841 (1): 25-27.
TL: Orenburg (S Ural, Volga distr., Russian Federation).

Genus Muschampia Tutt, 1906

ORIG: Muschampia Tutt, 1906. Nat. Hist. Brit. Lepid. 8: 218.
TS: Papilio proto Esper, 1830.

Subgenus Muschampia Tutt, 1906

ORIG: Muschampia Tutt, 1906. Nat. Hist. Brit. Lepid.8: 218.
TS: Papilio proto Esper, 1830.

Muschampia tessellum (Hübner, [1803])

ORIG: Papilio tessellum Hübner, [1803]. Samml. eur. Schmett. 1: pl. 93, fig. 469-470, [1806]: 70.
TL: Russian Federation.

Muschampia proto (Esper, [1808])

ORIG: Papilio proto Esper, [1808]. Schmett. Abb. Nat. Suppl. 2: pl. 123 (cont. 78), fig. 5-6, [1830]: 32-33.
TL: Portugal.

Muschampia proteides (Wagner, 1929)

ORIG: Hesperia proto ssp. proteides Wagner, 1929. Mitt. Münch. Entomol. Ges. 19(1): Erklärung zu Tafel II, pl. 2, fig. 26.
TL: Akşehir (Konya prov., Turkey).

Muschampia alta (Schwingenschuss, 1942)

ORIG: Hesperia proto ssp. alta Schwingenschuss, 1942. Z. Wien. Entomol.-Ver. 27(8): 182.
TL: Taormina (Messina prov., Sicily, Italy).
Taxonomic note. Muschampia alta Schwingenschuss, 1942, comb. & stat. nov. in: Hinojosa & al., 2021. Zool. J. Linn. Soc. 193: 854-855.

Muschampia mohammed (Oberthür, 1887)

ORIG: Syrichthus mohammed Oberthür, 1887. Bull. séances bibliogr. Soc. entomol. Fr. 1887: xlviii-xlix.
TL: Sebdou (Tlemcen prov., Algeria).

Muschampia leuzeae (Oberthür, 1881)

ORIG: Syrichthus leuzeae Oberthür, 1881. Étud. Entomol. 6: 60, pl. 3, fig. 10.
TL: Mascara (NW Algeria).

Subgenus Reverdinus Ragusa, [1919]

ORIG: Reverdinus Ragusa, [1919]. Naturalista sicil. 23(7/12): 172.
TS: Papilio altheae Hübner, 1800 (= Hesperia floccifera Zeller, 1847).

Muschampia floccifera (Zeller, 1847)

ORIG: Hesperia floccifera Zeller, 1847. Isis von Oken 1847(4): 286-287.
TL: Syracuse (Sicily, Italy).

Muschampia orientalis (Reverdin, 1913)

ORIG: Carcharodus orientalis Reverdin, 1913. Bull. Soc. lépid. Genève 2(1-4): 225, 230-235, pl. 21, fig. 14.
TL: Peloponnese (S Greece).

Muschampia stauderi (Reverdin, 1913)

ORIG: Carcharodus stauderi Reverdin, 1913. Bull. Soc. lépid. Genève 2(1-4): 225-230, pl. 21, fig. 12.
TL: El Kantara (Biskra prov., Algeria).

Muschampia baeticus (Rambur, [1839])

ORIG: Syrichtus baeticus Rambur, [1839]. Faune entomol. Andal. 2: pl. 12, fig. 3-4.
TL: Andalusia (S Spain).

Muschampia lavatherae (Esper, [1783])

ORIG: Papilio lavatherae Esper, [1783]. Schmett. Abb. Nat. 1(2): 148-149, pl. 82 (cont. 32), fig. 4.
TL: France ; Switzerland.

Genus Carcharodus Hübner, [1819]

ORIG: Carcharodus Hübner, [1819]. Verz. bekannt. Schmett.: 110.
TS: Papilio alceae Esper, 1780.

Carcharodus alceae (Esper, [1780])

ORIG: Papilio alceae Esper, [1780]. Schmett. Abb. Nat.1(2): 4-6, pl. 51 (cont. 1), fig. 3.
TL: Germany.

Carcharodus tripolina (Verity, 1925)

ORIG: Erynnis alceae race tripolina Verity, 1925. Entomol. Rec. J. Var. 37(4): 54.
TL: Garian plateau, S Tripoli (Lybia).

Tribe Erynnini Brues & Melander, 1932

ORIG: Erynninae Brues & Melander, 1932. Bull. Mus. comp. Zool. 73: 235-236.
TG: Erynnis Schrank, 1801.

Genus Erynnis Schrank, 1801

ORIG: Erynnis Schrank, 1801. Fauna Boica 2(1): 152, 157-160.
TS: Papilio tages Linnaeus, 1758.

Erynnis tages (Linnaeus, 1758)

ORIG: Papilio tages Linnaeus, 1758. Syst. Nat. (ed. 10)1: 485.
TL: Sweden.

Erynnis marloyi (Boisduval, [1834])

ORIG: Thanaos marloyi Boisduval, [1834]. Icon. Hist. Lépid. 1: 241, pl. 47, fig. 6-7.
TL: Peloponnese (S Greece).

Tribe Pyrgini Burmeister, 1878

ORIG: Pyrgidae Burmeister, 1878. Descr. phys. Rép. Argentine 5: 245-246.
TG: Pyrgus Hübner, [1819].

Genus Pyrgus Hübner, [1819]

ORIG: Pyrgus Hübner, [1819]. Verz. bekannt. Schmett.: 109.
TS: Papilio alveolus Hübner, 1800 (= Papilio malvae Linnaeus, 1758).

Subgenus Pyrgus Hübner, [1819]

ORIG: Pyrgus Hübner, [1819]. Verz. bekannt. Schmett.: 109.
TS: Papilio alveolus Hübner, 1800 (= Papilio malvae Linnaeus, 1758).

Pyrgus malvoides (Elwes & Edwards, 1897)

ORIG: Hesperia malvoides Elwes & Edwards, 1897. Trans. Zool. Soc. Lond. 14(4): 156, 160, pl. 23, fig. 27-27a [genitalia].
TL: Biarritz (SW France).

Pyrgus malvae (Linnaeus, 1758)

ORIG: Papilio malvae Linnaeus, 1758. Syst. Nat. (ed. 10)1: 485.
TL: Sweden.

Pyrgus melotis (Duponchel, [1834])

ORIG: Hesperia melotis Duponchel, [1834]. In: Godart, Hist. nat. Lépid. suppl. 1: 257-258, pl. 42, fig. 1-2.
TL: Lebanon.

Subgenus Scelotrix Rambur, 1858

ORIG: Scelotrix Rambur, 1858. Cat. syst. lépid. Andal.: 63-65.
TS: Papilio carthami Hübner, 1813.

Pyrgus carthami (Hübner, [1813])

ORIG: Papilio carthami Hübner, [1813]. Samml. eur. Schmett. 1: pl. 143, fig. 720-723.
TL: S Germany.

Pyrgus sidae (Esper, [1784])

ORIG: Papilio sidae Esper, [1784]. Schmett. Abb. Nat. 1(2): 178-179, pl. 90 (cont. 40), fig. 3.
TL: Volga region (Russian Federation).

Pyrgus centaureae (Rambur, [1839])

ORIG: Hesperia centaureae Rambur, [1839]. Faune entomol. Andal.2: pl. 8, fig. 10, [1840]: 315-316 (nota).
TL: Dovre (Norway).

Pyrgus cacaliae (Rambur, [1839])

ORIG: Hesperia cacaliae Rambur, [1839]. Faune entomol. Andal.2: pl. 8, fig. 6-7, k, [1840]: 313-314 (nota).
TL: Alps (SE France).

Pyrgus andromedae (Wallengren, 1853)

ORIG: Syrichtus andromedae Wallengren, 1853. Öfversigt Kongl. Vetenskaps-akad. förhandl. 10(2): 25.
TL: Dovre (Norway).

Subgenus Ateleomorpha Warren, 1926

ORIG: Pyrgus (Ateleomorpha) Warren, 1926. Trans. R. Entomol. Soc. London 74(1): 19, 87.
TS: Hesperia onopordi Rambur, [1839].

Pyrgus serratulae (Rambur, [1839])

ORIG: Hesperia serratulae Rambur, [1839]. Faune entomol. Andal. 2: pl. 8, fig. 9, m, [1840]: 318-319.
TL: Spain.

Pyrgus armoricanus (Oberthür, 1910)

Syrichthus alveus f. [sp., race?] armoricanus Oberthür, 1910. Étud. Lépid. comp. 4: 411-413, 681, pl. 57, fig. 509-517.
TL: Rennes (C France).

Pyrgus alveus (Hübner, [1803])

ORIG: Papilio alveus Hübner, [1803]. Samml. eur. Schmett. 1: pl. 92, fig. 461-463, [1806]: 70.
TL: Germany.

Pyrgus warrenensis (Verity, 1928)

ORIG: Hesperia alveus race warrenensis Verity, 1928. Bull. Soc. entomol. Fr. 1928(8): 140-141.
TL: Sulden (South Tyrol, Italy).

Pyrgus bellieri (Oberthür, 1910)

ORIG: Syrichthus alveus f. [sp., race?] bellieri Oberthür, 1910. Étud. Lépid. comp. 4: 404-406, 680, pl. 56, fig. 490-491.
TL: Larche (Alpes-de-Haute-Provence dep., SE France).

Pyrgus numida (Oberthür, 1910)

ORIG: Syrichthus alveus f. [sp., race?] numida Oberthür, 1910. Étud. Lépid. comp. 4: 404-405, 680, pl. 55, fig. 484-486.
TL: Tazoult (NE Algeria).

Pyrgus onopordi (Rambur, [1839])

ORIG: Hesperia onopordi Rambur, [1839]. Faune entomol. Andal. 2: pl. 8, fig. 13, p, [1840]: 319-320.
TL: Granada (Andalusia, S Spain).

Pyrgus carlinae (Rambur, [1839])

ORIG: Hesperia carlinae Rambur, [1839]. Faune entomol. Andal. 2: pl. 8, fig. 11, n, [1840]: 314-315 (nota).
TL: Alps (France ; Switzerland).

Pyrgus cirsii (Rambur, [1839])

ORIG: Hesperia cirsii Rambur, [1839]. Faune entomol. Andal. 2: pl. 8, fig. 12, o, [1840]: 315 (nota).
TL: Fontainebleau (C France).

Pyrgus cinarae (Rambur, [1839])

ORIG: Hesperia cinarae Rambur, [1839]. Faune entomol. Andal. 2: pl. 8, fig. 4-5, j.
TL: Sarepta (Volga reg., Russian Federation).