DYN CL SYN Subfamily Heteropterinae


		Subfamily Heteropterinae Aurivillius, 1925

Synonyms
[ORIG]	Heteropterinae Aurivillius, 1925. In: Seitz, Gross-Schmet. Erde 13: 506, 546.
		TG: Heteropterus Duméril, 1806.
[IN]	Eumesiidae Felder & Felder, [1867]. Reise Öst. Fregatte Novara, Zool. 2(2): 504 (invalid by ICZN, opinion 1669).
		TG: Eumesia Felder & Felder, [1867] (invalid by ICZN, opinion 1669).
[?]	Cyclopidinae Speyer, 1879. Entomol. Ztg. 40(10-12): 486.
		TG: Cyclopides Hübner, [1819] (= Heteropterus Duméril, 1806).
[JS]	Carterocephalini Orfila, 1949. Acta Zool. Lilloana 8: 584.
		TG: Carterocephalus Lederer, 1852.

Notes
(1)	The Subfamily name Eumesiidae Felder & Felder, [1867] was placed on the Official Index of Rejected and Invalid Family-group Names in Zoology by Opinion 1669 (ICZN, 1992. Bull. zool. Nomencl. 49(1): 88-89).
(2)	Verity, 1940 (Farf. diurne Ital. 1: 86) replaced Cyclopidinae Speyer, 1879 with Heteropterinae Aurivillius, 1925 because the type-genus name Cyclopides Hübner, [1819] is a junior synonym of Heteropterus Duméril, 1806 (ICZN Code, Article 40.2 provides that such action taken before 1961, and becoming widely accepted (as it has), is valid). However, in accordance with Recommendation 40A of the Code, the new name (Heteropterinae) should be cited with its author (Aurivillius) and date (1925), followed in parentheses by the date of priority of the name it replaces (1879).
(3)	List of the tribe found in the Western Palearctic: Heteropterini Aurivillius, 1925.
(4)	List of the tribe not found in the Western Palearctic: -.

Bibliography
Aurivillius C. 1925. 9. Familie: Hesperidae. In: Seitz, Die Gross-Schmetterlinge der Erde. 13. Band, Die afrikanischen Tagfalter. Text-Band. Stuttgart [Germany]: Alfred Kernen (Ed.). pp. 505-588. (url)
	New taxon: Heteropterinae.

Felder C. von- & Felder R. 1864-1867. Reise der Österreichischen Fregatte Novara um die Erde, Zoologischer Theil. Zweiter Band ; Zweite Abtheilung: Lepidoptera, Rhopalocera. Wien: Kaiserlich-Königlichen Hof- und Staatsdruckerei [Ed.]. pp. 1-548. (url)
	New taxon: Eumesiidae.

ICZN. 1992. Opinion 1669. Dalla Mabille, 1904 (Insecta, Lepidoptera): conserved. The Bulletin of zoological nomenclature 49(1): 88-89. (url)

Orfila R. N. 1949. Notas sistemáticas sobre Lepidoptera Rhopalocera. Acta Zoológica Lilloana 8: 583-586. (url not available)
	New taxon: Carterocephalini.

Speyer A. 1879. Die Hesperiden-Gattungen des europäischen Faunengebiets. II. Nachträge. Das Flügelgeäder. Entomologische Zeitung 40(10-12): 477-500. (url)
	New taxon: Cyclopidinae.

Verity R. 1940. Le Farfalle diurne d'Italia. Volume primo, Considerazioni generali e Superfamiglia Hesperides. Firenze: Marzocco SA (Ed.). pp. i-xxxiv, 1-131, pl. 1-4, pl. [morphology] I-II. (url not available)

Reference publications
Chiba H., Bozano G. C. & Fan X. 2023. Guide to the Butterflies of the palearctic region. Hesperiidae, part 1: Subfam. Coeliadinae & Heteropterinae. Milano: Omnes Artes (Ed.). pp. 1-71, fig. (url not available).

Toussaint E. F. A., Breinholt J. W., Earl C., Warren A. D., Brower A. V. Z., Yago M., Dexter K. M., Espeland M., Pierce N. E., Lohman D. J. & Kawahara A. Y. 2018. Anchored phylogenomics illuminates the skipper butterfly tree of life. BMC Evolutionary Biology 18(101): 1-11, fig. (https://doi.org/10.1186/s12862-018-1216-z)

Toussaint E. F. A, Ellis E. A., Gott R. J., Warren A. D., Dexter K. M., Storer C., Lohman D. J. &Kawahara A. Y. 2021. Historical biogeography of Heteropterinae skippers via Beringian and post-Tethyan corridors. Zoologica Scripta 50(1): 100-111, fig. (https://doi.org/10.1111/zsc.12457)

Warren A. D., Ogawa J. R. & Brower A. V. Z. 2009. Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic Entomology 34: 467–523, fig. (https://doi.org/10.1111/j.1365-3113.2008.00463.x)

Archives of Western Palearctic Lepidoptera
Editors-in-Chief: Michel Taymans & Sylvain Cuvelier